Coefficient of Conservatism:
Coefficient of Wetness:
B. S. Walters
An herbaceous vine in rich dense or dry open forests, fencerows, and thickets; often along pond and marsh borders, river bottoms, and railroads.
Al-Shehbaz & Schubert (1989) consider D. villosa to be a primarily Coastal Plain species with strongly ridged stems and leaves all alternate, while recognizing our plant as D. quaternata J. F. Gmel., with essentially terete stems and the lowest leaves on the stem whorled. Many, however, regard this variation as part of a broadly defined D. villosa. Herbarium collections often do not have the lower portions of the stem, but our plants in the field may have the lowermost leaves alternate or whorled in the same population, and the stems seem slightly ridged. Hopefully further work will clarify this situation.