Coefficient of Conservatism:
Coefficient of Wetness:
BEACH GRASS, MARRAM GRASS
A. A. Reznicek
Common to abundant on the sandy shores and dunes of Lakes Michigan, Huron, and Superior. May occur a short distance inland, especially near Lake Michigan, on older dunes and blowouts.
The only truly inland occurrence, entirely divorced from the modern Great Lakes, is on the University of Michigan Biological Station grounds on the southeast shore of Douglas Lake, Cheboygan Co. (first collected by J. H. Ehlers 7277 in 1920, MICH – and observed as recently as 2018). Its occurrence there is a modest mystery, though Solidago houghtonii, also long known from the same locality, was noted as having been planted there in 1935. Some other somewhat inland occurrences, e.g., in Muskegon Co., have spread from plantings to stabilize bare sand.
Observations on A. breviligulata at the Sleeping Bear Dunes, Leelanau Co., by F. C. Gates showed that the rhizomes may grow as much as 2.4 m in a single year as sand accumulates. In possessing both vertical and horizontal rhizomes, this plant is unusually well adapted for survival on sand dunes.
×Calammophila don-hensonii Reznicek & Judz. is a hybrid with Calamagrostis canadensis known from Lake Superior. See Delisle-Oldham, Oldham, & Catling (2008) for a morphological analysis of non-maritime Ammophila populations.