Coefficient of Conservatism:
Coefficient of Wetness:
CRESTED SHIELD FERN
Deciduous, mixed conifer-hardwood, or coniferous swamps; wet thickets, hummocks in bogs, fens, and sedge meadows; stream banks, lake shores, and seeps.
Dryopteris ×boottii (Tuck.) Underw. (D. cristata × D. intermedia), usually found in swamp forests, is one of the more frequent and easily recognized fern hybrids in Michigan. It is reminiscent of a rather narrow (and less cut) D. carthusiana, but is noticeably glandular.